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The Basilica of Saint Mark is a 13th-century Venetian church on the western edge of Lion Square, now repurposed as the Municipal Art Gallery of Heraklion. Originally the cathedral of the Duke of Crete, its elegant stone exterior and single-nave interior reflect the peak of Venetian ecclesiastical architecture on the island.
The basilica was built in 1239 by the Venetians and dedicated to Saint Mark, the patron saint of Venice. It was damaged by earthquakes in 1303 and 1508 and each time rebuilt. Under Ottoman rule it was converted into a mosque (the Defterdar Mosque). After Crete's union with Greece, it was restored and in 1956 became the Municipal Art Gallery.